macrocyclic

adj

1. Of an organic compound: having a closed ring of more than about twelve atoms.
2. Of or relating to a macrocycle.
3. Pertaining to a rust fungus whose life cycle includes all five spore stages: pycniospore (spermatium), aeciospore, urediniospore, teliospore, and basidiospore.

   “Almost all the rusts occurring on the ferns are both heteroecious and macrocyclic; so also are most of those occurring on the conifers.”